International Journal of Aquaculture, 2026, Vol.16, No.4, 213-233 http://www.aquapublisher.com/index.php/ija 231 Acknowledgments The auther would like to express their sincere gratitude to Dr. Zhang from Jiyang College, Zhejiang A&F University, for his valuable guidance and helpful suggestions during the preparation of this manuscript. Conflict of Interest Disclosure The auther affirms that this research was conducted without any commercial or financial relationships that could be construed as a potential conflict of interest. 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